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Insectivorous Plants

Page 312 of 400, printed 312

[page 362] Pinguicula or Nepenthes, for these plants are not at all closely related to the Droceraceae. But the difficulty is not nearly so great as it at first appears. Firstly, the juices of many plants contain an acid, and, apparently, any acid serves for digestion. Secondly, as Dr. Hooker has remarked in relation to the present subject in his address at Belfast (1874), and as Sachs repeatedly insists,* the embryos of some plants secrete a fluid which dissolves albuminous substances out of the endosperm; although the endosperm is not actually united with, only in contact with, the embryo. All plants, moreover, have the power of dissolving albuminous or proteid substances, such as protoplasm, chlorophyll, gluten, aleurone, and of carrying them from one part to other parts of their tissues. This must be effected by a solvent, probably consisting of a ferment together with an acid.** Now, in the case of plants which are able to absorb already soluble matter from captured insects, though not capable of true digestion, the solvent just referred to, which must be occasionally present in the glands, would be apt to exude from the glands together with the viscid secretion, inasmuch as endosmose is accompanied by exosmose. If such exudation did ever occur, the solvent would act on the animal matter contained within the captured insects, and this would be an act of true digestion. As it cannot be doubted that this process would be of high service to plants * Traite de Botanique 3rd edit. 1874, p. 844. See also for following facts pp. 64, 76, 828, 831. ** Since this sentence was written, I have received a paper by Gorup-Besanez ( Berichte der Deutschen Chem. Gesellschaft, Berlin, 1874, p. 1478), who, with the aid of Dr. H. Will, has actually made the discovery that the seeds of the vetch contain a ferment, which, when extracted by glycerine, dissolves albuminous substances, such as fibrin, and converts them into true peptones. [page 363] growing in very poor soil, it would tend to be perfected through natural selection. Therefore, any ordinary plant having viscid glands, which occasionally caught insects, might thus be converted under favourable circumstances into a species capable of true digestion. It ceases, therefore, to be any great mystery how several genera of 312
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