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The Descent of Man, and Selection in Relation to Sex

Page 538 of 825, printed 538

much more serious wound than several diverging ones. In Sir Philip Egerton s museum there is a horn of the red-deer (Cervus elaphus), thirty inches in length, with not fewer than fifteen snags or branches ; and at Moritzburg there is still preserved a pair of antlers of a red-deer, shot in 1699 by Frederick I., one of which bears the astonishing number of thirty-three branches and the other twenty-seven, making altogether sixty branches. Richardson figures a pair of antlers of the wild reindeer with twenty-nine points. (24. On the horns of red-deer, Owen, British Fossil Mammals, 1846, p. 478; Richardson on the horns of the reindeer, Fauna Bor. Americana, 1829, p. 240. I am indebted to Prof. Victor Carus, for the Moritzburg case.) From the manner in which the horns are branched, and more especially from deer being known occasionally to fight together by kicking with their fore-feet (25. Hon. J.D. Caton ( Ottawa Acad. of Nat. Science, May 1868, p. 9) says that the American deer fight with their fore-feet, after the question of superiority has been once settled and acknowledged in the herd. Bailly, Sur l Usage des cornes, Annales des Sciences Nat. tom. ii. 1824, p. 371.), M. Bailly actually comes to the conclusion that their horns are more injurious than useful to them. But this author overlooks the pitched battles between rival males. As I felt much perplexed about the use or advantage of the branches, I applied to Mr. McNeill of Colonsay, who has long and carefully observed the habits of red-deer, and he informs me that he has never seen some of the branches brought into use, but that the brow antlers, from inclining downwards, are a great protection to the forehead, and their points are likewise used in attack. Sir Philip Egerton also informs me both as to red-deer and fallow-deer that, in fighting, they suddenly dash together, and getting their horns fixed against each other s bodies, a desperate struggle ensues. When one is at last forced to yield and turn round, the victor endeavours to plunge his brow antlers into his defeated foe. It thus appears that the upper branches are used chiefly or exclusively for pushing and fencing. Nevertheless in some species the upper branches are used as weapons of offence; when a man was attacked by a wapiti deer (Cervus canadensis) in Judge Caton s park in Ottawa, and several men tried to rescue him, the stag never raised his head from the ground; in fact he kept his face almost flat on the ground, with his nose nearly between his fore feet, except when he rolled his head to one side to take a new observation preparatory to a plunge. In this position the ends of the horns were directed against his adversaries. In rolling his head he necessarily raised it 538
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