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The Descent of Man, and Selection in Relation to Sex

Page 393 of 825, printed 393

The season of love is that of battle; but the males of some birds, as of the game-fowl and ruff, and even the young males of the wild turkey and grouse (18. Audubon, Ornithological Biography, vol. ii. p. 492; vol. i. pp. 4-13.), are ready to fight whenever they meet. The presence of the female is the teterrima belli causa. The Bengali baboos make the pretty little males of the amadavat (Estrelda amandava) fight together by placing three small cages in a row, with a female in the middle; after a little time the two males are turned loose, and immediately a desperate battle ensues. (19. Mr. Blyth, Land and Water, 1867, p. 212.) When many males congregate at the same appointed spot and fight together, as in the case of grouse and various other birds, they are generally attended by the females (20. Richardson on Tetrao umbellus, Fauna Bor. Amer.: Birds, 1831, p. 343. L. Lloyd, Game Birds of Sweden, 1867, pp. 22, 79, on the capercailzie and black-cock. Brehm, however, asserts ( Thierleben, B. iv. s. 352) that in Germany the grey-hens do not generally attend the Balzen of the black-cocks, but this is an exception to the common rule; possibly the hens may lie hidden in the surrounding bushes, as is known to be the case with the gray-hens in Scandinavia, and with other species in N. America.), which afterwards pair with the victorious combatants. But in some cases the pairing precedes instead of succeeding the combat: thus according to Audubon (21. Ornithological Biography, vol. ii. p. 275.), several males of the Virginian goat-sucker (Caprimulgus virgianus) court, in a highly entertaining manner the female, and no sooner has she made her choice, than her approved gives chase to all intruders, and drives them beyond his dominions. Generally the males try to drive away or kill their rivals before they pair. It does not, however, appear that the females invariably prefer the victorious males. I have indeed been assured by Dr. W. Kovalevsky that the female capercailzie sometimes steals away with a young male who has not dared to enter the arena with the older cocks, in the same manner as occasionally happens with the does of the red-deer in Scotland. When two males contend in presence of a single female, the victor, no doubt, commonly gains his desire; but some of these battles are caused by wandering males trying to distract the peace of an already mated pair. (22. Brehm, Thierleben, etc., B. iv. 1867, p. 990. Audubon, Ornithological Biography, vol. ii. p. 492.) Even with the most pugnacious species it is probable that the pairing 393
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