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The Descent of Man, and Selection in Relation to Sex

Page 260 of 825, printed 260

Canfield for information with respect to the horns of the female: see also his paper in Proceedings of the Zoological Society, 1866, p. 109. Also Owen, Anatomy of Vertebrates, vol. iii. p. 627), only a few of the females, about one in five, have horns, and these are in a rudimentary state, though sometimes above four inches long: so that as far as concerns the possession of horns by the males alone, this species is in an intermediate condition, and the horns do not appear until about five or six months after birth. Therefore in comparison with what little we know of the development of the horns in other antelopes, and from what we do know with respect to the horns of deer, cattle, etc., those of the prong-horned antelope appear at an intermediate period of life, that is, not very early, as in cattle and sheep, nor very late, as in the larger deer and antelopes. The horns of sheep, goats, and cattle, which are well developed in both sexes, though not quite equal in size, can be felt, or even seen, at birth or soon afterwards. (41. I have been assured that the horns of the sheep in North Wales can always be felt, and are sometimes even an inch in length, at birth. Youatt says ( Cattle, 1834, p. 277), that the prominence of the frontal bone in cattle penetrates the cutis at birth, and that the horny matter is soon formed over it.) Our rule, however, seems to fail in some breeds of sheep, for instance merinos, in which the rams alone are horned; for I cannot find on enquiry (42. I am greatly indebted to Prof. Victor Carus for having made enquiries for me, from the highest authorities, with respect to the merino sheep of Saxony. On the Guinea coast of Africa there is, however, a breed of sheep in which, as with merinos, the rams alone bear horns; and Mr. Winwood Reade informs me that in one case observed by him, a young ram, born on Feb. 10th, first shewed horns on March 6th, so that in this instance, in conformity with rule, the development of the horns occurred at a later period of life than in Welsh sheep, in which both sexes are horned.), that the horns are developed later in life in this breed than in ordinary sheep in which both sexes are horned. But with domesticated sheep the presence or absence of horns is not a firmly fixed character; for a certain proportion of the merino ewes bear small horns, and some of the rams are hornless; and in most breeds hornless ewes are occasionally produced. Dr. W. Marshall has lately made a special study of the protuberances so common on the heads of birds (43. Uber die knochernen Schadelhocker der Vogel, in the Niederland. Archiv fur Zoologie, B.i. Heft 2, 260
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