northern parts, where the land almost joins, and where, under a
slightly different climate, there might have been free migration for
the northern temperate forms, as there now is for the strictly arctic
productions. We see the same fact in the great difference between the
inhabitants of Australia, Africa, and South America under the same
latitude: for these countries are almost as much isolated from each
other as is possible. On each continent, also, we see the same fact;
for on the opposite sides of
lofty and continuous mountain-ranges, and of great deserts, and
sometimes even of large rivers, we find different productions; though
as mountain chains, deserts, etc., are not as impassable, or likely to
have endured so long as the oceans separating continents, the
differences are very inferior in degree to those characteristic of
distinct continents.
Turning to the sea, we find the same law. No two marine faunas are more
distinct, with hardly a fish, shell, or crab in common, than those of
the eastern and western shores of South and Central America; yet these
great faunas are separated only by the narrow, but impassable, isthmus
of Panama. Westward of the shores of America, a wide space of open
ocean extends, with not an island as a halting-place for emigrants;
here we have a barrier of another kind, and as soon as this is passed
we meet in the eastern islands of the Pacific, with another and totally
distinct fauna. So that here three marine faunas range far northward
and southward, in parallel lines not far from each other, under
corresponding climates; but from being separated from each other by
impassable barriers, either of land or open sea, they are wholly
distinct. On the other hand, proceeding still further westward from the
eastern islands of the tropical parts of the Pacific, we encounter no
impassable barriers, and we have innumerable islands as halting-places,
until after travelling over a hemisphere we come to the shores of
Africa; and over this vast space we meet with no well-defined and
distinct marine faunas. Although hardly one shell, crab or fish is
common to the above-named three approximate faunas of Eastern and
Western America and the eastern Pacific islands, yet many fish range
from the Pacific into the Indian Ocean, and many shells are common to
the eastern islands of the Pacific
and the eastern shores of Africa, on almost exactly opposite meridians
of longitude.
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