On the absence of intermediate varieties at the present day. On the
nature of extinct intermediate varieties; on their number. On the vast
lapse of time, as inferred from the rate of deposition and of
denudation. On the poorness of our pal ontological collections. On the
intermittence of geological formations. On the absence of intermediate
varieties in any one formation. On the sudden appearance of groups of
species. On their sudden appearance in the lowest known fossiliferous
strata.
In the sixth chapter I enumerated the chief objections which might be
justly urged against the views maintained in this volume. Most of them
have now been discussed. One, namely the distinctness of specific
forms, and their not being blended together by innumerable transitional
links, is a very obvious difficulty. I assigned reasons why such links
do not commonly occur at the present day, under the circumstances
apparently most favourable for their presence, namely on an extensive
and continuous area with graduated physical conditions. I endeavoured
to show, that the life of each species depends in a more important
manner on the presence of other already defined organic forms, than on
climate; and, therefore, that the really governing conditions of life
do not graduate away quite insensibly like heat or moisture. I
endeavoured, also, to show that intermediate varieties, from existing
in lesser numbers than the forms which they connect, will generally be
beaten out and exterminated during the course of further modification
and improvement. The main cause, however, of innumerable intermediate
links not now occurring everywhere throughout nature depends
on the very process of natural selection, through which new varieties
continually take the places of and exterminate their parent-forms. But
just in proportion as this process of extermination has acted on an
enormous scale, so must the number of intermediate varieties, which
have formerly existed on the earth, be truly enormous. Why then is not
every geological formation and every stratum full of such intermediate
links? Geology assuredly does not reveal any such finely graduated
organic chain; and this, perhaps, is the most obvious and gravest
objection which can be urged against my theory. The explanation lies,
as I believe, in the extreme imperfection of the geological record.
In the first place it should always be borne in mind what sort of
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